Subordinate Taxa
Scientific Name:
Lophozonia Turcz., Bull. Soc. Imp. Naturalistes Moscou 31: 396 (1858)
  • Nothofagus subgen. Lophozonia (Turcz.) Krasser, Ann. K. K. Naturhist. Hofmus. 11: 162 (1896)
  • = Nothofagus subgen. Nothofagus sect. Calusparassus subsect. Menziesiae Philipson & M.N.Philipson, Bot. J. Linn. Soc. 98: 34 (1988)
  • = Nothofagus subgen. Menziesospora R.S.Hill & J.Read, Bot. J. Linn. Soc. 105: 69 (1991)
  • = Nothofagus subgen. Nothofagus sect. Adenofagus Baum.-Bod., Syst. Fl. Neu-Caledonien 94, 114 (1992)
Type Taxon:
Lophozonia heterocarpa Turcz. = Lophozonia obliqua (Mirb.) Heenan & Smissen

Evergreen trees up to 40 m high. Stipules non-peltate. Leaves distichous, margin crenate; fimbrial vein complete; large, globular, glandular trichomes present on cuticle; solitary unicellular trichome type A absent; solitary unicellular trichomes type C present, with a large base equal to or greater than the diameter of the trichome and with a large, unicellular, thin-walled trichome emerging. Stomata randomly oriented; without thickened T-pieces of cuticle at the poles separating the two guard cells; without giant stomata over the major veins; stomata size within the areoles variable. Upper epidermal cells over veins not distinguishable from areolar cells, without granular cell walls. Male dichasium a solitary pseudanthium; perianth open and broadly campanulate, asymmetric and irregular; stamens usually >20, developing centrifugally; anthers often slightly curved; distal connective protrusion weakly developed, filament connective free, without epidermal papillae or with isomorphic rounded papillae. Female dichasium with 1 central dimerous flower and 2 lateral trimerous flowers, or 2 trimerous flowers; stigmas clavate and weakly decurrent. Cupule valves 4, with simple trichomes; with rows of stalked appendages, hairy with globular, glandular apices; nuts triquetrous or lenticular, 2- or 3-winged.

Pollen: Peritreme or goniotreme, mesocolpia straight to convex; colpi long, tenuimarginate, with V-shaped ends, margins not thickened, aperture >15.0 µm long, polar to equatorial length ratio 0.35–0.4.


A genus of seven species in Australia, New Zealand, Chile and Argentina. Lophozonia has two endemic species in Australia, four endemic species in southern South America (Chile and Argentina) and one endemic species in New Zealand.

Indigenous (Non-endemic)
Number of species in New Zealand within Lophozonia Turcz.
Indigenous (Endemic)1
Baumann-Bodenheim, M.G. 1992: Trisyngyne. Systematik der Flora von Neu-Caledonien (Melanesien-Südpazifik). Mrs A. L. Baumann, Herrliberg.
Du Rietz, G.E. 1930: On domatia in the genus Nothofagus . Svensk Botanisk Tidskrift 24: 4.
Heenan, P.B.; Smissen, R.D. 2013: Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146(1): 1–31.
Hill, R.S.; Read, J. 1991: A revised infrageneric classification of Nothofagus (Fagaceae). Botanical Journal of the Linnean Society 105: 37–72.
Krasser, F. 1896: Bemerkungen zur Systematik der Buchen. Annalen des K. K. Naturhistorischen Hofmuseums 11: 155–163.
Ogden, J. 1989: On the coenospecies concept and tree migrations during the oscillations of the Pleistocene climate. Journal of the Royal Society of New Zealand 19: 249–262.
Ogden, J.; Stewart, G.H.; Allen, R.B. 1996: Ecology of New Zealand Nothofagus forests. In: Veblen, T.T.; Hill, R.S.; Read, J. (ed.) The ecology and biogeography of Nothofagus forests. Yale University Press, New Haven. 82–25.
Peterson, K.R.; Pfister, D.H.; Bell, C.D. 2010: Cophylogeny and biogeography of the fungal parasite Cyttaria and its host Nothofagus, southern beech. Mycologia 102: 1417–1425.
Philipson, W.R.A.; Philipson, M.N. 1988: A classification of the genus Nothofagus (Fabaceae). Botanical Journal of the Linnean Society 98: 27–36.
Poole, A.L. 1950a: Studies of New Zealand Nothofagus 1. Taxonomy and Floral Morphology. Transactions of the Royal Society of New Zealand 78: 323–380.
Poole, A.L. 1950b: Studies of the New Zealand Nothofagus species 2. Nut and cupule development. Transactions of the Royal Society of New Zealand 78(4): 502–508.
Poole, A.L. 1952: The development of Nothofagus seed. Transactions of the Royal Society of New Zealand 80: 207–212.
Poole, A.L. 1987: Southern Beeches. Science Information Publishing Centre, DSIR, Wellington.
Rozefelds, A.C.; Drinnan, A.N. 2002: Ontogeny of pistillate flowers and inflorescences in Nothofagus subgenus Lophozonia (Nothofagaceae). Plant Systematics and Evolution 233: 105–126.
Turczaninow, C.G.J. 1858: Animadversiones in secundam partem herbarii Turczaninowiani, nunc Universitatis Caesareae Charkowiensis. Bulletin de la Société Imperiale des Naturalistes de Moscou. Moscow 31: 380–500.
Wardle, J. 1984: The New Zealand Beeches. Ecology, Utilisation and Management. New Zealand Forest Service. Caxton Press, Christchurch, New Zealand.
Wilcox, M.D.; Ledgard, N.J. 1983: Provenance variation in the New Zealand species of Nothofagus. New Zealand Journal of Ecology 6: 19–31.