Classification
 Nomenclature
Scientific Name:
Veronica hectorii Hook.f., Handb. New Zealand Fl. 212 (1864) – as hectori
Synonymy:
  • Hebe hectorii (Hook.f.) Cockayne & Allan, Trans. New Zealand Inst. 57: 40 (1926)
  • Leonohebe hectorii (Hook.f.) Heads, Bot. Soc. Otago Newsl. 5: 8 (1987)
Lectotype (designated by Ashwin, in Allan 1961): Otago, Mt Alta, Hector no. 27, 1863, Herb. Hookerianum, K (small broken piece, mounted on lower right of sheet that also includes material collected by Sinclair and Haast, and Hector and Buchanan)
Etymology:
The epithet hectorii honours James Hector (1834–1907), geologist, explorer, and founder of several scientific institutions in New Zealand.
Vernacular Name(s):
whipcord hebe
 Description

Whipcord shrub to 1.0 m tall. Stems ascending to erect, glabrous except for a line of eglandular hairs at the connate leaf bases. Leaf bud indistinct, its outer leaves fully grown, diverging. Leaves opposite-decussate, connate in pairs and encircling stem, appressed and usually covering the well-marked node above, scale-like; lamina coriaceous, deltoid to broadly deltoid to sub-orbicular, 1.5–3.5 mm long, 2.0–4.5 mm wide, glossy green, olive-green, or yellowish-green above and beneath; midrib and veins not evident; surfaces glabrous; margin ciliolate, entire (incised to dentate in juvenile and reversion leaves); apex obtuse, sub-acute, apiculate, or mucronate; base broad; petiole absent. Inflorescence a terminal spike, 3.5–15.0 mm long; flowers crowded, 4–16, all bisexual or some female; bracts opposite-decussate, connate, deltoid; pedicels absent. Calyx lobes 4–5 (usually free or rarely anterior pair fused from ⅓ to ⅔ of the way to apex; 5th lobe small, posterior), obtuse to sub-acute, sub-equal, 1.5–3.0 mm long, usually eglandular-ciliate with long, deflexed, sinuous hairs, sometimes ciliolate, mixed with short glandular hairs as well. Corolla 5.0–8.5 mm diameter, tube white, 1.5–2.5 mm long, ≤calyx, eglandular-hairy inside; lobes 4, white, erect to spreading, sub-equal, oblong to ovate to broadly elliptic, 3.0–4.5 mm long, obtuse; nectar guides absent. Stamen filaments white, 3.5–4.2 mm long; anthers pink to purple. Style glabrous, 3.3–6.5 mm long. Capsule latiseptate, obtuse to sub-acute, 1.8–3.2 mm long, 1.8–2.5 mm at widest point. Seeds ellipsoid to oblong, flattened, smooth, straw-yellow or brown, 0.9–1.4 mm long.

 Key
1Leaf apex of some or most leaves apiculate or mucronatesubsp. demissa
Leaf apex sub-acute or obtuse or rounded2
2Leaves longer than to about as long as broad, mostly 1.2–2.0 mm, occasionally to 2.7 mm, long (northern South Island)subsp. coarctata
Leaves broader than long, mostly 1.7–3.1 mm, occasionally 1.2–2.7 mm long (southern South Island)subsp. hectorii
 Recognition

The differences between V. hectorii and V. tetragona plants are slight and were discussed in detail by Bayly & Kellow (2006), along with discussion of various options for taxonomic treatment at species rank. Although they are very similar, differences in flavonoid chemistry and distribution support their recognition at species rank (Bayly & Kellow 2006).

V. tetragona and V. hectorii can be reliably distinguished by the thickened leaf apex of V. tetragona, compared to that of V. hectorii, which is not thickened (Bayly & Kellow 2006, Fig. 48, p. 91). The strongly tetragonous to cruciform leafy branchlets of V. tetragona subsp. tetragona provide a clear distinction from V. hectorii, but the more rounded branchlets of subsp. subsimilis are not so easily distinguished, particularly from V. hectorii subsp. coarctata.

Bracts and calyx lobes, but not the leaves, often have one to several longitudinal ribs, resembling the leaves of V. lycopodioides and V. poppelwellii.

 Distribution

South Island: Western Nelson, Sounds Nelson, Westland, Canterbury, Otago Southland, Fiordland.

Stewart I.

 Habitat

Montane to alpine grassland and shrubland.

 Biostatus
Indigenous (Endemic)
Number of subspecific taxa in New Zealand within Veronica hectorii Hook.f.
CategoryNumber
Indigenous (Endemic)3
Total3
 Hybridisation

Veronica hectorii hybridises occasionally with V. odora, to form whipcord-like plants, but with longer leaves and internodes. In general, the parentage of V. odora × whipcord hybrids is hard to deduce from morphology, and in the field proximity of putative parents is the best criterion.

 Phenology

Flowers: November–April; Fruit: January–June (persisting all year).

 Cytology

2n = 40 (Bayly & Kellow 2006, as Hebe hectorii).

 Notes

Veronica hectorii is classified in V. subg. Pseudoveronica sect. Hebe and informally in the “Flagriformes” group, also known as whipcord hebes (Albach & Meudt 2010; Bayly & Kellow 2006).

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
Cockayne, L.; Allan, H.H. 1926: The present taxonomic status of the New Zealand species of Hebe. Transactions of the New Zealand Institute 57: 11–47.
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.
Hooker, J.D. 1864: Handbook of the New Zealand Flora: a systematic description of the native plants of New Zealand and the Chatham, Kermadec's, Lord Auckland's, Campbell's and Macquarie's Islands. Part I. Reeve, London.
Wagstaff, S.J.; Wardle, P. 1999: Whipcord Hebes - systematics, distribution, ecology and evolution. New Zealand Journal of Botany 37(1): 17–39.