Classification
 Nomenclature
Scientific Name:
Veronica cupressoides Hook.f., Handb. New Zealand Fl. 212 (1864)
Synonymy:
  • Hebe cupressoides (Hook.f.) Andersen, Trans. New Zealand Inst. 56: 693 (1926)
  • Leonohebe cupressoides (Hook.f.) Heads, Bot. Soc. Otago Newsl. 5: 8 (1987)
Lectotype (first designated by Moore, in Allan 1961; designated by Bayly & Kellow 2006 more precisely): Lindis Pass & Lake Dis[trict], on river flats, Hector and Buchanan no. 7, K (sprig in lower left corner of sheet only)
Etymology:
Cupressoides, cypress-like (genus Cupressus), a reference to the habit of the plants.
Vernacular Name(s):
cypress koromiko; cypress-like hebe; whipcord hebe
 Description

Whipcord shrub to 2 m tall. Stems erect, glabrous or eglandular-pubescent; hairs bifarious. Leaf bud indistinct, its outer leaves fully grown, diverging. Leaves opposite-decussate, connate in pairs and encircling stem, scale-like, appressed (slightly spreading when dry); lamina sub-coriaceous, ovate to deltoid, 0.8–2.0 mm long; 0.4–2.0 mm wide, dull glaucescent to yellowish-green above and beneath; veins not evident; surfaces glabrous or sometimes with dense, minute, glandular hairs; margin eglandular- or glandular-ciliolate, entire; apex sub-acute to obtuse; bases broad; petiole absent. Inflorescence a terminal spike, 3–37 mm long; flowers crowded, 2–22, all bisexual; bracts opposite-decussate, connate, ovate to deltoid; pedicels absent. Calyx lobes 4, seemingly 2–3 because posterior pair completely fused, anterior pair fused ⅔ or more, these pairs fused together to about halfway, obtuse to rounded, 0.5–1.0 mm long, eglandular-ciliate or -ciliolate mixed with short glandular cilia. Corolla 3–4 mm diameter; tube white to purplish, 0.9–1.4 mm long, = calyx, eglandular-hairy inside; lobes 4, white, pink, pale blue or pale purplish, sub-erect to recurved, sub-equal, elliptic, oblong, or sub-orbicular, 2–3 mm long, obtuse; nectar guides absent. Stamen filaments white, 2.1–4.0 mm long, anthers pink, magenta, or brownish. Style glabrous, 2–3 mm long. Capsule angustiseptate, emarginate, glabrous, 1.9–2.4 mm long, 0.9–1.4 mm at widest point. Seeds ovoid, ellipsoid, oblong, or obovoid, weakly flattened, smooth, pale brown, 0.7–1.1 mm long.

 Recognition

Veronica cupressoides is a distinctive species with a unique combination of characters: a whipcord growth form with tiny appressed, scale-like, dull to glaucescent leaves; long internodes (two to three times the leaves) with conspicuous nodal joints where the leaves are attached; the calyx lobes all fused for about half their length, the posterior pair completely fused, and the anterior pair fused ⅔ or more; narrow elliptic or oblong sub-equal corolla lobes; and angustiseptate capsules. Other unique features are the nectarial disc covering the lower third of the ovary at flowering and the aromatic foliage.

 Distribution

South Island: Marlborough (south-west), Canterbury (eastern foothills), and Otago (western parts, south to Wye Creek).

 Habitat

Montane, in scrub, rock outcrops, moraine, and slips, often near lakes and streams. Recorded elevations range from 300 to 1372 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: December–February; fruits: February–May (persisting all year).

 Cytology

2n = 42 (see Bayly & Kellow 2006, as Leonohebe cupressoides).

 Notes

Veronica cupressoides is classified in V. subg. Pseudoveronica sect. Hebe and informally in the “semi-whipcord hebe” group (Albach & Meudt 2010). It is sister to the other semi-whipcord hebes, which are smaller subshrubs with similar, rather glaucous leaves but short internodes.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Andersen, J. 1926: Popular names of New Zealand plants. Transactions of the New Zealand Institute 56: 659–714.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Leonohebe cupressoides (Hook.f.) Heads] [Nationally Endangered]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe cupressoides (Hook.f.) Cockayne & Allan]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Nationally Endangered]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Leonohebe cupressoides (Hook.f.) Heads] [Nationally Endangered]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.
Hooker, J.D. 1864: Handbook of the New Zealand Flora: a systematic description of the native plants of New Zealand and the Chatham, Kermadec's, Lord Auckland's, Campbell's and Macquarie's Islands. Part I. Reeve, London.