Classification
 Nomenclature
Scientific Name:
Veronica lavaudiana Raoul, Choix Pl. Nouv.-Zél. p. 16, t. 10 (1846)
Synonymy:
  • Hebe lavaudiana (Raoul) Cockayne & Allan, Trans. New Zealand Inst. 57: 44 (1926)
  • Parahebe lavaudiana (Raoul) Heads, Bot. J. Linn. Soc. 115: 83 (1994)
  • Heliohebe lavaudiana (Raoul) Garn.-Jones, New Zealand J. Bot. 34: 427 (1996)
Lectotype (designated by Garnock-Jones 1993): Nouvelle Zelande-Presqu'lle de Banks, Raoul, 1843, top left-hand piece on the sheet, this piece is recognisable as the left-hand branch of the piece illustrated in the protologue, P. Isolectotypes: K, MEL 1595921
Etymology:
"I have dedicated this pretty species to Monsieur Lavaud, Captain of the vessel, as a token of recognition for all the facilities and the friendly support which he has kindly consented to accord me in my scientific researches in New Zealand" (Raoul 1846, translated).
 Description

Low shrub to 0.3 m tall. Stems prostrate to ascending, eglandular-pubescent, or sometimes glandular as well; hairs uniform. Leaf bud indistinct; leaves separating while very small, opposite-decussate, spreading to recurved, separating early, coriaceous, obovate to orbicular, 7–30 mm long, 6–17 mm wide, dull dark green to glaucescent above, paler beneath; midrib evident, glabrous; margin glabrous, or glandular- and eglandular-ciliate, crenate or crenate-serrate with 6–10 pairs of teeth; apex obtuse to truncate, sometimes apiculate; base cuneate; petiole 2–8 mm long. Inflorescence a terminal compound spike, 20–150 mm long; flowers crowded, 50–150 per inflorescence, all bisexual; bracts alternate, the lowest opposite, deltoid to narrowly deltoid, = calyx; pedicels absent. Calyx lobes 5, acuminate, equal, 4–5 mm long, sometimes the anterior pair fused ⅔ of length, glandular-ciliate. Corolla 11–13 mm diameter; tube white with greenish-yellow base,1.5–2.0 mm long, < calyx, glabrous; lobes 4, white or pink (pink in bud) erect to spreading, unequal, elliptic to rhomboid or broadly rhomboid, 4.5–5.5 mm long, sub-acute; nectar guides absent. Stamen filaments white, 2.5 mm long; anthers yellow. Style glabrous, 6–7 mm long. Capsules broadly angustiseptate to turgid, emarginate, glabrous or glandular-hairy at apex, 3–4 mm long, 2.0–2.5 mm at widest point. Seeds obovoid or fusiform, weakly flattened, winged, weakly rugulose on back, pale brown, 1.7–2.7 mm long.

 Recognition

V. lavaudiana belongs to the distinctive sun hebe group, which is characterised by toothed or crenate leaves, terminal paniculate inflorescences, style very much > stamens, ciliate nectarial disc, and fusiform seeds. Within this grouping it is distinguished by its dull, crenate leaves, crowded white flowers with pink buds, acuminate calyx lobes, and glandular-hairy inflorescences. The leaves often have red margins.

 Habitat

Basalt outcrops and cliffs, generally confined to places out of reach of browsing mammals. Recorded elevations range from 152 to 809 m.

 Biostatus
Indigenous (Endemic)
 Hybridisation

V. lavaudiana does not grow together with any closely related species, but hybridises in cultivation with V. hulkeana (=V. ×fairfieldii Hook.f., Garnock-Jones 2008).

 Phenology

Flowers: October–January; fruits: January–March.

 Cytology

2n = 42 (Hair 1967, as Hebe lavaudiana).

 Notes

Veronica lavaudiana is classified in V. subg. Pseudoveronica sect. Hebe and informally in the “sun hebe” group (Albach & Meudt 2010). Monophyly of the sun hebe clade is firmly established (Albach & Meudt 2010), and this small clade is probably related to the speedwell hebe clade (formerly Parahebe p.p. and V. macrantha).

The leaves have dense stomata on both surfaces, but these are absent from the margins. The rounded teeth each have a sunken hydathode at the tip.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington. [as Hebe lavaudiana (Raoul) Anderson; Heliohebe lavaudiana (Raoul) Garn.-Jones]
Cockayne, L.; Allan, H.H. 1926: The present taxonomic status of the New Zealand species of Hebe. Transactions of the New Zealand Institute 57: 11–47.
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Heliohebe lavaudiana (Raoul) Garn.-Jones] [Declining]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Heliohebe lavaudiana (Raoul) Garn.-Jones]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Declining]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Heliohebe lavaudiana (Raoul) Garn.-Jones] [Declining]
Garnock-Jones, P.J. 1993: Heliohebe (Scrophulariaceae-Veroniceae), a new genus segregated from Hebe. New Zealand Journal of Botany 31: 323–339.
Garnock-Jones, P.J. 1996: Addendum. New Zealand Journal of Botany 34: 427.
Garnock-Jones, P.J. 2008: Botanical names for hybrids in Veronica sect. Hebe (Plantaginaceae) from New Zealand. New Zealand Journal of Botany 46: 523–529.
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Garnock-Jones, P.J.; Lloyd, D.G. 2004: A taxonomic revision of Parahebe (Plantaginaceae) in New Zealand. New Zealand Journal of Botany 42: 181–232. [as Heliohebe lavaudiana (Raoul) Garn.-Jones; Parahebe lavaudiana (Raoul) Heads]
Hair, J.B. 1967: Contributions to a chromosome atlas of the New Zealand flora — 10. Hebe (Scrophulariaceae). New Zealand Journal of Botany 5: 322–352.
Heads, M.J. 1994: A biogeographic review of Parahebe (Scrophulariaceae). Botanical Journal of the Linnean Society 115: 65–89.
Raoul, E. 1846: Choix de plantes de la Nouvelle-Zélande: recueillies et décrites. Fortin Masson, Paris.