Classification
 Nomenclature
Scientific Name:
Veronica mooreae (Heads) Garn.-Jones in Garnock-Jones et al., Taxon 56: 578 (2007)
Synonymy:
  • Leonohebe mooreae Heads, Bot. Soc. Otago Newsl. 5: 10 (1987)
  • Hebe mooreae (Heads) Garn.-Jones, Austral. Syst. Bot. 6: 479 (1993)
Holotype: Douglas Range, South Westland, 4100’, in low scrub on a steep gully side, P. Wardle, 16 Dec 1978, CHR 321236
  • = Leonohebe mooreae var. telmata Heads, Bot. Soc. Otago Newsl. 5: 10 (1987)
Holotype: Douglas Range, South Westland, 3500’, abundant in Chionochloa grassland on rolling country, P. Wardle, 16 Dec 1978, CHR 231327
Etymology:
Named after prominent New Zealand botanist Lucy Beatrice Moore (1906–1987), who prepared the treatment of Hebe (except the whipcord spp.) for Flora of New Zealand Vol. 1 (Allan 1961).
 Description

Shrub to 1.2 m tall or rarely to 2.0 m. Stems erect, eglandular-pubescent, hairs bifarious. Leaf bud distinct, its leaves appressed at margins until fully grown; sinus shield-shaped to rhomboid, or sometimes narrow and acute. Leaves opposite-decussate, erecto-patent; lamina coriaceous, rigid, oblong, oblong-elliptic, oblong-lanceolate, or linear-oblong, 7–28 mm long, 3–8 mm wide, glossy dark green above, somewhat glossy green to dark green beneath; midrib and  two lateral veins evident; surfaces glabrous; margin glabrous, entire but minutely crenulate; apex acute or sub-acute; base steeply cuneate to truncate; petiole 1–3 mm long. Inflorescence a lateral spike or raceme, 8–30 mm long; flowers crowded, 3–13, all bisexual; bracts opposite-decussate, connate, ovate or deltoid, > pedicels; pedicels erect, 0–1 mm long, sparsely eglandular-pubescent all around. Calyx lobes 4, anterior pair sometimes fused in lower ⅓, obtuse to sub-acute, 3–4 mm long, sub-equal, eglandular-ciliolate or mixed glandular- and eglandular-ciliolate. Corolla 10–12 mm diameter; tube white, 4–5 mm long, ≥ calyx; glabrous or with mixed eglandular and glandular hairs inside; lobes 4, white, erecto-patent to recurved, sub-equal, elliptic to broadly elliptic, 5–6 mm long, obtuse to rounded; nectar guides absent. Stamen filaments white, 3–4 mm long; anthers pink or pale pink. Style glabrous, 5.5–8.5 mm long. Capsules latiseptate, acute or sub-acute, glabrous, 3.5–4.5 mm long, 2.0–2.8 mm at widest point.  Seeds broadly ellipsoid, flattened, smooth, pale brown, 1.2–1.8 mm long.

 Recognition
Morphological features to distinguish Veronica odora and look-alikes. Species are ordered according to similarity of their habit.
 

odora

mooreae

pauciramosa

masoniae

venustula

brachysiphon

Leaf bud sinus

broad, shield-shaped

narrow & acute to broad & shield shaped

broad, shield-shaped

broad, shield-shaped

narrow, acute

narrow, acute

Leaf margin

sharply bevelled; glabrous

bevelled at 90º to surfaces, glabrous

rounded; glabrous or with minute hairs or denticles

rounded, papillate towards apex; ciliolate when young

weakly bevelled; ciliolate when young, becoming glabrous or papillate

weakly bevelled; ciliolate to ciliate when young, becoming papillate

Stomata

adaxial – (but often + at Arthur’s Pass); abaxial +

adaxial – (+ at Caswell Sound, Denniston); abaxial +

adaxial +; abaxial +

adaxial +; abaxial +

adaxial ±; abaxial +

adaxial ±; abaxial +

Midrib

sharply keeled beneath

depressed above; prominent beneath

rounded beneath and flattened just short of apex

keeled throughout

evident but not keeled

evident but not keeled

Inflorescence

terminal + usually lateral spikes

lateral spikes only

lateral spikes only

terminal spikes only

lateral raceme, sometimes ternate, rarely compound.

lateral raceme, sometimes ternate.

Bracts

not overtopping calyx

< calyx

< calyx

≥ calyx

≥ pedicels, < calyx

≥ pedicels, < calyx

Bracts and flowers

opposite

opposite

opposite

opposite

opposite below, becoming alternate

alternate, or lowermost opposite

Pedicels

0 mm

0–1 mm

0–0.5 mm

0 mm

0.5–7.0  mm

0.6–3.0 mm

Calyx, anterior lobes

free

free or fused to ⅓-way

fused > ⅔-way

free

free

free

Corolla lobes

narrow

broad

narrow

broad

± broad

± broad

 Distribution

South Island: Western Nelson (western mountains, Gouland Downs), Westland (Denniston Plateau and south of Mt Wilberg), Fiordland, Southland (Hump Ridge and Longwood Range).

Records from Canterbury (Wild Mans Brother Range, Cameron River, Laing, WELT17242) and Otago (East Dome, CHR 4138; regarded as anomalous by Bayly & Kellow 2006) are not mapped here.

 Habitat

Penalpine grassland and montane to sub-alpine scrub. Recorded elevations range from 500 to 1385 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: November–February, sometimes to June); fruits: January–June (sometimes persisting to be present all year).

 Cytology

2n = 126 (see Bayly & Kellow 2006, as Hebe mooreae).

 Notes

Veronica mooreae is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Buxifoliatae” (Albach & Meudt 2010; Bayly & Kellow 2006). The phylogenetic relationships of V. mooreae are not well resolved. On the basis of ITS sequence data, it falls within a large, unresolved grouping of shrubby hebes that have 2n = 40, 80, 120, not close to V. odora (2n = 42, 84), with which it was previously confused. Its unusual chromosome number  (2n = 126) suggests it might have a hybrid origin, and it is possible that one of its parent species is V. odora.

The thickened leaf margins are usually bevelled at about 90° to the surfaces, and their minute crenulations are visible with a lens or can be felt by rubbing a fingernail along the margin.

Plants from Gouland Downs have long internodes and slender leaves that are almost linear-elliptic.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington. [as Hebe mooreae (Heads) Garn.-Jones; Leonohebe mooreae Heads var. mooreae]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Not Threatened]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe mooreae (Heads) Garn.-Jones] [Not Threatened]
Garnock-Jones, P.J. 1993: Phylogeny of the Hebe complex (Scrophulariaceae: Veroniceae). Australian Systematic Botany 6(5): 457–479.
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.