Classification
 Nomenclature
Scientific Name:
Veronica ochracea (Ashwin) Garn.-Jones in Garnock-Jones et al., Taxon 56: 578 (2007)
Synonymy:
  • Hebe ochracea Ashwin in Allan, Fl. New Zealand 1, 936 (1961)
  • Leonohebe ochracea (Ashwin) Heads, Bot. Soc. Otago Newsl. 5: 7 (1987)
Holotype: Cobb Valley, N. W. Nelson, F. G. Gibbs, CHR 97077. Isotype: AK 8243 [according to Ashwin (in Allan 1961), although details on labels differ slightly]
Etymology:
The epithet ochracea is a reference to the yellowish to brownish colour of the leafy stems.
Vernacular Name(s):
whipcord hebe
 Description

Spreading low or bushy whipcord shrub to 0.45 m tall. Stems ascending or spreading, with short, erect secondary branches increasing in length away from main stem apex, eglandular-pubescent; hairs bifarious above leaf axils and usually in the deep groove between the connate leaf bases. Leaf bud indistinct, its outer leaves fully grown, diverging. Leaves opposite-decussate, appressed but not usually covering the obscure node above, crowded and overlapping, coriaceous, broadly ovate to rhomboid, 1.0–1.5 mm long, 1.5–3.0 mm wide, glossy yellow- to bronze-green above and beneath; veins not evident; surfaces glabrous; margin shortly ciliate or ciliolate, entire; apex obtuse; base broad, connate in pairs and encircling stem; petiole absent. Inflorescence a terminal spike, 2.0–8.5 mm long; flowers crowded, 4–8, all bisexual; bracts opposite-decussate and connate, broadly ovate; pedicels absent. Calyx lobes 4, seemingly 3 because anterior pair fused to apex, obtuse to sub-acute, 1.5–2.3 mm long, unequal, eglandular-ciliate with a few short glandular hairs as well. Corolla 4.5–7.5 mm diameter; tube white, 1.0–1.5 mm long, ≤ calyx, eglandular-hairy inside; lobes 4, white, sub-erect to spreading, unequal, elliptic to oblong, 3–4 mm long, obtuse to rounded; nectar guides absent. Stamen filaments white, 3.5–4.5 mm long; anthers pink or yellowish-pink. Style glabrous or a few bristles at base, 3–4 mm long. Capsules latiseptate, obtuse, glabrous or a few bristles especially along suture and at apex, 2.2–3.3 mm long, 1.7–2.2 mm at widest point. Seeds ellipsoid to obovoid, flattened, smooth, pale brown, 1.2–1.5 mm long.

 Recognition

Plants of V. ochracea and three other whipcord hebes, V. annulata, V. armstrongii, and V. salicornioides, all share fused anterior calyx lobes and exposed but generally obscure nodal joint. They all have allopatric distributions, with V. ochracea being confined to north-west Nelson and northern Westland. V. ochracea differs from all of them in its unusual chromosome number. Distinguishing morphological characters are relatively indistinct and subjective, the best being the yellowish- to bronze-green of the foliage in V. ochracea. The leaves are not as closely appressed as those of V. salicornioides and internodes are generally a little shorter. In dried specimens the leaves spread a little from the stem, but not as much as in plants of V. annulata and V. armstrongii.

 Distribution

South Island: Western Nelson (Anatoki Range, Cobb Valley and Peel Range, Mt Arthur Range, Owen Range), Mt Buckland, Paparoa Range.

 Habitat

Sub-alpine grassland and scrub, usually on calcareous substrates. Recorded elevations range from 915 to 1800 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: November–February; fruits: February–April, persisting longer.

 Cytology

2n = 124 (see Bayly & Kellow 2006, as Hebe ochracea).

 Notes

Veronica ochracea is classified in V. subg. Pseudoveronica sect. Hebe and informally in the “Flagriformes” group, also known as “whipcord hebes” (Albach & Meudt 2010; Bayly & Kellow 2006).

The sexual system needs to be checked in wild populations of V. ochracea. Some herbarium specimens (e.g., WELT 17567, 17481, 17563) seem to have small flowers and pollen was not seen in their dehisced anthers. Others have large flowers with copious pollen (e.g., WELT 82795).

Bayly and Kellow did not record seed characters, and the seed description above is based on only one specimen (WELT SP090750, Lake Peel, Sneddon). Seeds might be more variable than suggested here.

Morphology and chromosome numbers indicate a relationship with V. annulata, V. armstrongii, and V. salicornioides. Its unusual and high chromosome number suggests the possibility of a hybrid origin involving a species with 2n = 40, such as V. hectorii, and one with 2n = 84, such as V. armstrongii. However, V. armstrongii and V. ochracea are geographically separated.

Cultivars

​​​​​​​V. ochracea is common in cultivation as hebe ‘James Stirling’. The cv is said to be a dwarf form but otherwise no different from wild plants.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington. [as Hebe ochracea Ashwin; Leonohebe ochracea (Ashwin) Heads]
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe ochracea Ashwin] [Naturally uncommon]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe ochracea Ashwin]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Naturally Uncommon]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe ochracea Ashwin] [Naturally Uncommon]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.