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Classification
Nomenclature
Scientific Name:
Veronica perbella (de Lange) Garn.-Jones in Garnock-Jones et al., Taxon 56: 578 (2007)
Synonymy:
- ≡ Hebe perbella de Lange, New Zealand J. Bot. 36: 399 (1998)
Holotype: New Zealand, Northland, Tutamoe Ecological District, Waima Forest, Hauturu High Point Track, P. J. de Lange 3169 & I. McFadden, 7 Nov 1996, AK 230119. Isotypes: CHR 487605, K, WELT 79994
Etymology:
Vernacular Name(s):
Bartlett’s hebe
Description
Bushy shrub to 1.8 m tall. Stems erect, glabrous. Leaf bud distinct, its leaves appressed until fully grown; sinus absent. Leaves opposite-decussate, erecto-patent to spreading; lamina coriaceous, elliptic or narrowly oblanceolate, 35–110 mm long, 7–23 mm wide, glossy dark green above, dull pinkish-green beneath; midrib and faint pinnate secondary veins evident; surfaces glabrous except for minute glandular and eglandular hairs along midrib above; margin glabrous, entire; apex acute to obtuse; base cuneate; petiole absent or indistinct, 1–2 mm long. Inflorescence a lateral raceme, 60–150 mm long; flowers more or less crowded, 15–80, all bisexual; bracts alternate to loosely whorled, lanceolate, < pedicels; pedicels spreading, 2–5 mm long, minutely puberulent all around. Calyx lobes 4 or occasionally small posterior 5th lobe present, narrowly acute to acuminate, 2.2–3.8 mm long, equal, with sparse, minute, sessile glandular hairs on face, mixed glandular- and eglandular-ciliolate. Corolla 5–9 mm diameter; tube white, 1.8–2.2 mm long, ≤ calyx, eglandular-puberulent inside; lobes 4, purplish, sometimes pink, red, or carmine, fading to whitish, erect or sub-erect, reflexed with age, unequal, lanceolate to narrowly ovate, 4.8–6.5 mm long, acute; nectar guides absent. Stamen filaments purplish, 7–10 mm long; anthers blue to purple. Style glabrous, 8–10 mm long. Capsules latiseptate, acute, glabrous except for a few short eglandular hairs in septal groove, 7–8 mm long, 5–6 mm at widest point. Seeds ovate, flattened, weakly papillate, pale brown, 1.8–2.0 mm long.
Recognition
adamsii | macrocarpa | perbella | saxicola | |
|---|---|---|---|---|
Stem hairs | uniformly to bifariously puberulent or glabrous | uniformly to bifariously puberulent | glabrous | usually glabrous or rarely bifarious-puberulent |
Leaf bud sinus | present | absent | absent | absent |
Lamina (above) | glossy | glossy | glossy | dull |
Calyx colour | dark green or bronze-green | yellow-green | pink, reddish, violet, or red- to brown-green | yellowish- to dark green |
Calyx lobes outer surface | minute sessile glands | glabrous or eglandular-puberulent | minute sessile glands | minute sessile glands |
Corolla colour | white to mauve, rarely violet | white, rarely lilac, sometimes tinged lilac or pink | violet-red, violet, pink, deep mauve, very rarely carmine | pale lavender (rarely lilac) |
Corolla tube | ≤ calyx; hairy inside | > calyx; hairy inside | ≤ calyx; hairy inside | < calyx; glabrous inside |
Corolla lobes | subacute | obtuse to rounded | acute | subacute or bluntly acute |
Capsule | glabrous | glabrous | glabrous except for a few short hairs along septal grooves | glabrous |
Distribution | far North, known only from four sites, all within 1 km2, west of Spirits Bay | southern Northland from Whangārei and southern Kaipara Harbour southwards to South Auckland, Coromandel, and near Kawhia | western Northland between Ahipara and Waima Forest | known only from Maungaraho Rock, near Dargaville |
Distribution
North Island: Northland (Ahipara, Herekino Forest, Warawara Forest, Waimā Forest). Formerly present also at Kaitaia (de Lange & Rolfe 2008).
Habitat
Rock outcrops and shallow soils in cloud forest, also in kauri forest and gum-land scrub. Recorded elevations range from 100 to 690 m.
Biostatus
Indigenous (Endemic)
Phenology
Flowers: March–December; fruits: May–March.
Cytology
2n = 40 (see Bayly & Kellow 2006, as Hebe perbella).
Notes
Veronica perbella is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Occlusae” (Albach & Meudt 2010; Bayly & Kellow 2006).
Images
Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
de Lange, P.J. 1998: Hebe perbella (Scrophulariaceae) - a new and threatened species from western Northland, North Island, New Zealand. New Zealand Journal of Botany 36(3): 399–406.
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe perbella de Lange] [Nationally Endangered]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe perbella de Lange]
de Lange, P.J.; Rolfe, J.R. 2008: Hebe saxicola (Plantaginaceae) - a new threatened species from western Northland, North Island, New Zealand. New Zealand Journal of Botany 46: 531–545.
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Nationally Endangered]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe perbella de Lange] [Nationally Endangered]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
