Classification
 Nomenclature
Scientific Name:
Veronica stricta Banks & Sol. ex Benth. in de Candolle, Prodr. 10 459 (1846) var. stricta
Synonymy:
  • = Veronica lindleyana Paxton, Paxton's Mag. Bot. 12: 247 (1846)
  • Veronica stricta var. lindleyana (Paxton) J.B.Armstr., Trans. New Zealand Inst. 13: 356 (1881)
  • = Veronica macroura Hook.f. ex Benth. in de Candolle, Prodr. 10 459 (1846)
  • Hebe macroura (Hook.f. ex Benth.) Cockayne & Allan, Trans. New Zealand Inst. 57: 15 (1926)
  • Hebe stricta var. macroura (Hook.f. ex Benth.) L.B.Moore in Allan, Fl. New Zealand 1, 906 (1961)
  • Veronica stricta var. macroura (Hook.f. ex Benth.) Garn.-Jones in Garnock-Jones et al., Taxon 56: 579 (2007)
Holotype: “Veronica macroura Hook fil”, N. Zealand, Herb. Benthamianum, K
  • = Veronica cookiana Colenso, Trans. & Proc. New Zealand Inst. 20: 201 (1888)
  • Veronica macroura var. cookiana (Colenso) Cheeseman, Man. New Zealand Fl. 501 (1906)
  • Hebe cookiana (Colenso) Cockayne & Allan, Trans. New Zealand Inst. 57: 16 (1926)
Lectotype (designated by Moore, in Allan 1961): Table Cape plant, H. Hill, 1887, Herb. Colenso, WELT 5315
  • = Veronica parkinsoniana Colenso, Trans. & Proc. New Zealand Inst. 21: 97 (1889)
  • Hebe parkinsoniana (Colenso) Cockayne, Trans. New Zealand Inst. 60: 472 (1929)
Type: none designated
  • = Veronica salicifolia var. atkinsonii Cockayne, Trans. New Zealand Inst. 48: 200 (1916)
  • Hebe salicifolia var. atkinsonii (Cockayne) Andersen, Trans. New Zealand Inst. 56: 693 (1926)
  • Hebe stricta var. atkinsonii (Cockayne) L.B.Moore in Allan, Fl. New Zealand 1, 906 (1961)
Type: none designated (see notes by Moore in Allan 1961)
  • = Veronica salicifolia var. longiracemosa Cockayne, Trans. & Proc. New Zealand Inst. 49: 61 (1917)
  • Hebe salicifolia var. longiracemosa (Cockayne) Cockayne & Allan, Trans. New Zealand Inst. 57: 18 (1926)
Lectotype (designated by Moore, in Allan 1961): on steep bank at outskirts of forest, Moumahaki (Egmont-Wanganui bot. distr. of N. Z.), lowland belt, L. Cockayne 8163, March 1916, K
  • = Hebe stricta (Banks & Sol. ex Benth.) L.B.Moore in Allan, Fl. New Zealand 1, 904 (1961) var. stricta
Vernacular Name(s):
kōkōmuka; kōkoromiko; kōkoromuka; korohiko; korokio; koromiko; koromuka
 Description

Stems brown to black when dry, glabrous to densely puberulent. Leaves lanceolate to elliptic to oblanceolate or obovate, 19–127 mm long, 5.5–45.0 mm wide; lamina thin to sub-coriaceous, usually puberulent to pubescent on midribs and margins, occasionally glabrous on midribs, rarely glabrous on margins. Calyx lobes pubescent (north of about Manawatu Gorge) or glabrous on faces, mixed glandular- and eglandular-ciliolate on margins; corolla tube ≥ lobes.

 Recognition

Veronica stricta var. stricta as circumscribed here is a variable complex of regional forms. Plants are large shrubs with glabrous to puberulent, rarely pubescent, stems, thin to sub-coriaceous leaves that are usually hairy along the midrib above and below and ciliolate along the margins.

Coastal plants throughout its range tend to have broader, more coriaceous leaves, often obovate. North of about Manawatu Gorge plants have hairs on the faces of the calyx lobes, whereas further south the calyx lobes are ciliolate on the margins only. Plants in Gisborne and East Cape tend to have very long inflorescences and may have glabrous stems.

In Coromandel and on Great Barrier I. (Aotea I.) some plants have longer hairs on the leaf midribs and margins and thus approach the appearance of V. pubescens. V. pubescens can usually be distinguished by having a small sinus in the leafy bud (but this is often obscured by dense hairs) and hairy corolla lobes (outside).

In the South Island, V. stricta var. stricta overlaps slightly in its distribution with the similar V. salicifolia. The most reliable distinction is the presence of a leaf bud sinus in V. salicifolia plants, absent in V. stricta plants.

 Distribution

North Island: throughout.

South Island: Western Nelson (near the coast of western Golden Bay), Sounds Nelson (near the coast), Marlborough (coastal), North Canterbury (near the coast north of Napenape).

 Habitat

Coastal to montane, usually in open sites: rock outcrops, banks, roadsides, scrub, forest clearings, slips, etc., and may be weedy in urban areas. Recorded elevations range from 0 to 1451 m.

 Biostatus
Indigenous (Endemic)
 Hybridisation

It is likely that V. stricta hybridises with other species, which could explain some of its variability. Binomial names believed to be based on hybrids are listed below:

Veronica ×affinis (Cheeseman) Garn.-Jones, New Zealand Journal of Botany 46: 524 (2008) ≡ ​​​​​​​Veronica macrocarpa var. affinis Cheeseman, Manual of the New Zealand Flora: 505 (1906). ≡ Hebe ×affinis (Cheeseman) Cockayne & Allan, Transactions of the New Zealand Institute 57: 20 (1926). The plants are considered to be V. macrocarpa × stricta. The illegitimate name Hebe ×macrosala also applies to this hybrid group (see Garnock-Jones 2008). Veronica macrocarpa × stricta (Garnock-Jones 2008).

Veronica ×carsei Petrie, Transactions of the New Zealand Institute 55: 96 (1924, as V. carsei). ≡ Hebe carsei (Petrie) Cockayne, Transactions of the New Zealand Institute 60: 469 (1929). = Hebe ×laevisala Cockayne & Allan, Transactions of the New Zealand Institute 57: 46 (1926). Thought to be Veronica stricta var. stricta × V. venustula (Garnock-Jones 2008).

Veronica ×simmonsii Cockayne, Transactions of the New Zealand Institute 48: 202 (1916). ≡ Hebe ×simmonsii (Cockayne) Cockayne & Allan, Transactions of the New Zealand Institute 56: 20 (1926). = ​​​​​​​Hebe ×angustisala Cockayne & Allan, Transactions of the New Zealand Institute 57: 46 (1926). Specimens are consistent with parentage (V. stenophylla × V. stricta var. atkinsonii) suggested in the protologue (Garnock-Jones 2008).

 Phenology

Flowers: January–July, extending to September; fruits: January–September, persisting all year.

 Cytology

2n = 40, 80 (see Bayly & Kellow 2006, as Hebe stricta var. stricta).

 Notes

Previous treatments (Moore, in Allan 1961; Bayly & Kellow 2006) have recognised further varieties that here are included under var. stricta. The patterns of variation are more complex than reliance on a few characters, such as sepal indumentum (var. atkinsonii) and leaf width (var. macroura), would indicate.

Cultivars

Veronica ×andersonii Lindley & Paxton, Paxton’s Flower Garden 2: t. 38 (1851, as V. andersonii). ≡ Hebe ×andersonii (Lindl. & Paxton) Cockayne, Transactions of the New Zealand Institute 60: 468 (1929). Likely to be V. speciosa × stricta (Heenan 1994). Heenan (1994) clarified the origins and naming of several cultivars of V. ×andersonii (as Hebe ×andersonii).

 Images
 Bibliography
Andersen, J. 1926: Popular names of New Zealand plants. Transactions of the New Zealand Institute 56: 659–714.
Armstrong, J.B. 1881: A synopsis of the New Zealand species of Veronica, Linn., with notes on new species. Transactions of the New Zealand Institute 13: 344–359.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington. [as Hebe cookiana (Colenso) Cockayne & Allan; Hebe macroura (Hook.f. ex Benth.) Cockayne & Allan; Hebe parkinsoniana (Colenso) Cockayne; Hebe salicifolia var. atkinsonii (Cockayne) Andersen; Hebe salicifolia var. atkinsonii (Cockayne) Cockayne & Allan; Hebe salicifolia var. longiracemosa (Cockayne) Cockayne & Allan; Hebe stricta (Banks & Sol. ex Benth.) L.B.Moore; Hebe stricta (Banks & Sol. ex Benth.) L.B.Moore var. stricta; Hebe stricta var. atkinsonii (Cockayne) L.B.Moore; Hebe stricta var. macroura (Hook.f. ex Benth.) L.B.Moore; Veronica cookiana Colenso; Veronica lindleyana Paxton; Veronica macroura Hook.f. ex Benth.; Veronica macroura var. cookiana (Colenso) Cheeseman; Veronica parkinsoniana Colenso; Veronica salicifolia var. atkinsonii Cockayne; Veronica salicifolia var. longiracemosa Cockayne; Veronica stricta var. lindleyana (Paxton) J.B.Armstr.]
Cheeseman, T.F. 1906: Manual of the New Zealand Flora. Government Printer, Wellington.
Cockayne, L. 1916: Notes on New Zealand floristic botany, including descriptions of new species, etc. (No. 1). Transactions of the New Zealand Institute 48: 193–202.
Cockayne, L. 1917: Notes on New Zealand floristic botany, including descriptions of new species & c. (No. 2). Transactions and Proceedings of the New Zealand Institute 49: 56–65.
Cockayne, L. 1929: New combinations in the genus Hebe. Transactions of the New Zealand Institute 60: 465–472.
Cockayne, L.; Allan, H.H. 1926: The present taxonomic status of the New Zealand species of Hebe. Transactions of the New Zealand Institute 57: 11–47.
Colenso, W. 1888: On new Phænogamic plants of New Zealand. Transactions and Proceedings of the New Zealand Institute 20: 188–211.
Colenso, W. 1889: A description of some newly-discovered Phænogamic plants; being a further contribution towards the making known the botany of New Zealand. Transactions and Proceedings of the New Zealand Institute 21: 80–108.
de Candolle, A.P. 1846: Prodromus Systematis Naturalis Regni Vegetabilis. Vol. 10. Treuttel et Würtz, Paris.
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Not Threatened; Taxonomically indistinct]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe stricta (Banks & Sol. ex Benth.) L.B.Moore var. stricta; Hebe stricta var. atkinsonii (Cockayne) L.B.Moore; Hebe stricta var. macroura (Hook.f. ex Benth.) L.B.Moore] [Not Threatened]
Garnock-Jones, P.J. 2008: Botanical names for hybrids in Veronica sect. Hebe (Plantaginaceae) from New Zealand. New Zealand Journal of Botany 46: 523–529.
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heenan, P.B. 1994: The origin and identification of Hebe ×andersonii and its cultivars (Scrophulariaceae). Horticulture in New Zealand, Journal of the Royal New Zealand Institute of Horticulture 5: 21–25.
Hooker, J.D. 1852–1853 ("1853"): The Botany of the Antarctic Voyage of H.M. Discovery Ships Erebus and Terror in the Years 1839–1843, under the command of Captain Sir James Clark Ross. II. Flora Novae-Zelandiae. Part I. Flowering plants. Lovell Reeve, London.
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