Classification
 Nomenclature
Scientific Name:
Veronica kellowiae Garn.-Jones in Garnock-Jones et al., Taxon 56: 577 (2007)
Synonymy:
  • Hebe ramosissima G.Simpson & J.S.Thomson, Trans. & Proc. Roy. Soc. New Zealand 72: 29 (1942)
  • Leonohebe ramosissima (G.Simpson & J.S.Thomson) Heads, Bot. Soc. Otago Newsl. 5: 7 (1987)
Lectotype (designated by Moore, in Allan 1961): Mount Tapuaenuku, moist debris at 2150 m alt., G. Simpson, CHR 75691. Isolectotype: AK 107861
Etymology:
The epithet kellowiae honours Alison Valerie Kellow, who revised the taxonomy of the “Connatae” to which the species belongs and was co-author of the 2006 revision of the shrubby hebes (Bayly & Kellow 2006).
 Description

Sub-shrub or spreading low shrub to 0.15 m tall. Stems decumbent, eglandular-pubescent; hairs bifarious. Leaf bud distinct, its outer leaves fully grown, diverging. Leaves opposite-decussate to sub-distichous, connate in pairs and encircling stem, erecto-patent when young, becoming spreading to recurved; lamina coriaceous, narrowly obovate, sometimes elliptic to obovate to rhomboid, 3.3–9.5 mm long, 1.5–5.5 mm wide, dull to somewhat glossy, green to dark green above, green beneath; midrib obscure, slightly thickened beneath; surfaces glabrous; margin glabrous or sparsely glandular- and eglandular-ciliolate, especially at base and petiole, entire; apex acute to rounded; base cuneate; petiole indistinct, 0.5–1.5 mm long. Inflorescence a terminal compound spike, 8–20 mm long; flowers crowded, 6–20 per inflorescence in terminal and 0–4 lateral spikes, female or bisexual on separate plants, ⚥ > ♀; bracts opposite-decussate and connate or free below, the lowermost sometimes sterile, becoming alternate above, elliptic, ovate, or oblong, < calyx; pedicels absent. Calyx lobes 4, sub-acute to obtuse, 2.0–3.5 mm long, unequal, with mixed, very short eglandular and minute glandular cilia. Corolla 3–6 mm diameter; tube white, 2.8–3.5 mm long, < calyx, glabrous; lobes 4, white, sub-erect to recurved, sub-equal, ovate, orbicular, or rhomboid, 3–4 mm long, obtuse; nectar guides absent. Stamen filaments white, 0.5–1.0 mm long; anthers magenta. Style glabrous, 2–4 mm long. Capsules latiseptate, acute, glabrous, 3.7–4.0 mm long, 1.8–3.0 mm at widest point. Seeds ellipsoid, flattened, smooth, pale brown, 0.7–0.8 mm long.

 Recognition

V. kellowiae plants are most similar to V. macrocalyx in growth form, leaf shape and overall appearance, but V. macrocalyx plants differ in having usually more spikes (up to 12) per inflorescence, and usually a longer calyx that exceeds the corolla tube. The few-flowered inflorescences also distinguish V. kellowiae from other “Connatae”. V. petriei plants also seem similar in their growth form and long corolla tubes, but differ in having leaves minutely puberulent on the adaxial surface near the base, spiralled bracts, many of them sterile at the base of the inflorescence, stalked flowers, and a longer calyx and corolla.

 Distribution

South Island: Marlborough (Inland Kaikōura Range); Canterbury (Seaward Kaikōura Range, Mt Terako).

 Habitat

Alpine rocks and scree, often in moist places. Recorded elevations range from 1200 to 2652 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: December–February; fruits: December–March.

 Cytology

2n = 42 (see Bayly & Kellow 2006, as Hebe ramosissima).

 Notes

Veronica kellowiae is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Connatae” (Albach & Meudt 2010; Bayly & Kellow 2006), related to V. epacridea, V. haastii, and V. macrocalyx, according to phylogenetic analyses of ITS sequence data (Wagstaff et al. 2002).

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe ramosissima G.Simpson & J.S.Thomson] [Naturally uncommon]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe ramosissima G.Simpson & J.S.Thomson]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Naturally Uncommon]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe ramosissima G.Simpson & J.S.Thomson] [Naturally Uncommon]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.
Kellow, A.V.; Bayly, M.J.; Mitchell, K.A.; Markham, K.R.; Brownsey, P.J. 2003: A taxonomic revision of Hebe informal group "Connateae" (Plantaginaceae), based on morphology and flavonoid chemistry. New Zealand Journal of Botany 41(4): 613–635. [as Hebe ramosissima G.Simpson & J.S.Thomson; Leonohebe ramosissima (G.Simpson & J.S.Thomson) Heads]
Simpson, G.; Thomson, J.S. 1942: Notes on some New Zealand plants and descriptions of new species (No. 2). Transactions and Proceedings of the Royal Society of New Zealand 72: 21–40.
Wagstaff, S.J.; Bayly, M.J.; Garnock-Jones, P.J.; Albach, D.C. 2002: Classification, origin, and diversification of the New Zealand Hebes (Scrophulariaceae). Annals of the Missouri Botanical Garden 89: 3–63.